<?xml version="1.0" encoding="ISO-8859-1"?><article xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance">
<front>
<journal-meta>
<journal-id>1816-7950</journal-id>
<journal-title><![CDATA[Water SA]]></journal-title>
<abbrev-journal-title><![CDATA[Water SA]]></abbrev-journal-title>
<issn>1816-7950</issn>
<publisher>
<publisher-name><![CDATA[Water Research Commission (WRC)]]></publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id>S1816-79502012000200004</article-id>
<title-group>
<article-title xml:lang="en"><![CDATA[Synthetic monthly flow duration curves for the Cape Floristic Region, South Africa]]></article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Hope]]></surname>
<given-names><![CDATA[Allen]]></given-names>
</name>
<xref ref-type="aff" rid="A01"/>
</contrib>
<contrib contrib-type="author">
<name>
<surname><![CDATA[Bart]]></surname>
<given-names><![CDATA[Ryan]]></given-names>
</name>
<xref ref-type="aff" rid="A01"/>
</contrib>
</contrib-group>
<aff id="A01">
<institution><![CDATA[,San Diego State University Department of Geography ]]></institution>
<addr-line><![CDATA[San Diego CA]]></addr-line>
<country>USA</country>
</aff>
<pub-date pub-type="pub">
<day>00</day>
<month>00</month>
<year>2012</year>
</pub-date>
<pub-date pub-type="epub">
<day>00</day>
<month>00</month>
<year>2012</year>
</pub-date>
<volume>38</volume>
<numero>2</numero>
<fpage>191</fpage>
<lpage>200</lpage>
<copyright-statement/>
<copyright-year/>
<self-uri xlink:href="http://www.scielo.org.za/scielo.php?script=sci_arttext&amp;pid=S1816-79502012000200004&amp;lng=en&amp;nrm=iso&amp;tlng=en"></self-uri><self-uri xlink:href="http://www.scielo.org.za/scielo.php?script=sci_abstract&amp;pid=S1816-79502012000200004&amp;lng=en&amp;nrm=iso&amp;tlng=en"></self-uri><self-uri xlink:href="http://www.scielo.org.za/scielo.php?script=sci_pdf&amp;pid=S1816-79502012000200004&amp;lng=en&amp;nrm=iso&amp;tlng=en"></self-uri><abstract abstract-type="short" xml:lang="en"><p><![CDATA[A flow duration curve (FDC) provides a valuable planning and management tool since it describes the entire flow regime of a river. Water resource planning in South Africa is often based on monthly river flow data and synthetic FDCs are required for applications in ungauged catchments. The objective of this study was to derive 11 monthly FDC percentile flows and the mean annual flow (MAQ) for catchments in the Cape Floristic Region of South Africa using regression equations with readily measureable catchment variables, including vegetation indices from Moderate Resolution Imaging Spectrometer (MODIS) satellite imagery. An 'all-models' approach with 10-fold validation was adopted to identify the 'best' regression models. Predictions of percentile flows above the median flow and MAQ were generally good but poor for low flows. Overall predictive uncertainty had a tendency to be larger in drier catchments. The most important predictive variables were catchment mean annual precipitation, physiography and soils. MODIS vegetation indices were significant predictors in equations for 6 percentile flows and MAQ, and predictive uncertainty increased if the MODIS indices were excluded from model development. The regression approach implemented in this study may be appropriate for other regionalisation studies that are based on a small sample of gauged catchments.]]></p></abstract>
<kwd-group>
<kwd lng="en"><![CDATA[Western Cape Region]]></kwd>
<kwd lng="en"><![CDATA[flow duration curve]]></kwd>
<kwd lng="en"><![CDATA[ungauged catchments]]></kwd>
<kwd lng="en"><![CDATA[multiple regression]]></kwd>
<kwd lng="en"><![CDATA[cross-validation]]></kwd>
</kwd-group>
</article-meta>
</front><body><![CDATA[ <p align="right"><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><b>ARTICLES</b></font></p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="4"><b><a name="top"></a>Synthetic    monthly flow duration curves for the Cape Floristic Region, South Africa</b></font></p>     <p>&nbsp;</p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><b>Allen Hope</b><a href="#back"><sup>*</sup></a><b>;</b>    <b> Ryan Bart</b></font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">Department of Geography,    San Diego State University, San Diego, CA 92182, USA</font></p>     <p>&nbsp;</p>     <p>&nbsp;</p> <hr size="1" noshade>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><b>ABSTRACT</b></font></p>     ]]></body>
<body><![CDATA[<p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">A flow duration    curve (FDC) provides a valuable planning and management tool since it describes    the entire flow regime of a river. Water resource planning in South Africa is    often based on monthly river flow data and synthetic FDCs are required for applications    in ungauged catchments. The objective of this study was to derive 11 monthly    FDC percentile flows and the mean annual flow (MAQ) for catchments in the Cape    Floristic Region of South Africa using regression equations with readily measureable    catchment variables, including vegetation indices from Moderate Resolution Imaging    Spectrometer (MODIS) satellite imagery. An 'all-models' approach with 10-fold    validation was adopted to identify the 'best' regression models. Predictions    of percentile flows above the median flow and MAQ were generally good but poor    for low flows. Overall predictive uncertainty had a tendency to be larger in    drier catchments. The most important predictive variables were catchment mean    annual precipitation, physiography and soils. MODIS vegetation indices were    significant predictors in equations for 6 percentile flows and MAQ, and predictive    uncertainty increased if the MODIS indices were excluded from model development.    The regression approach implemented in this study may be appropriate for other    regionalisation studies that are based on a small sample of gauged catchments.</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><b>Keywords:</b>    Western Cape Region, flow duration curve, ungauged catchments, multiple regression,    cross-validation</font></p> <hr size="1" noshade>     <p>&nbsp;</p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="3"><b>Introduction</b></font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">A flow duration    curve (FDC) is a graphical representation of the frequency distribution of the    complete river flow regime and is one of the most commonly-used techniques in    hydrology (Croker et al., 2003). While empirical FDCs can be developed using    gauged flow data, estimation of FDCs in ungauged catchments requires a regionalisation    approach which is usually based on flow information from a network of gauged    sites. The International Association of Hydrological Sciences (IAHS) Decade    on Predictions in Ungauged Basins (PUB) is an international initiative that    recognises the critical need to advance hydrological predictions in ungauged    catchments (Sivapalan, 2003). River flow prediction in ungauged catchments is    widely regarded as the ultimate challenge in hydrology (Sivapalan, 2003).</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">A number of regionalisation    approaches have been proposed for estimating FDCs in ungauged catchments. A    common regionalisation methodology describes the FDC in terms of a mathematical    model and then relates the parameters of the model to catchment morphological    and/or climatic variables using regression analysis (Niadas, 2005; Viola et    al., 2011). Probabilistic models can be used to describe the FDC and regression    models developed to estimate the parameters of the distribution (Castellarin    et al., 2007)</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">Assumptions regarding    models that describe the form of FDCs can be avoided by developing regional    regression equations to predict the selected percentile flows (e.g., flows for    exceedance percentages 5%, 10%, 20%, ...95%). Catchment physical characteristics    are used as the predictor variables in these regression equations (e.g., Mohamoud,    2008; Yu and Yang, 2000). This approach has been used successfully by Yu and    Yang (2000) in Taiwan to predict daily stream flow for 10 percentile flows in    ungauged catchments. Precipitation was largely uniform across the study region    and catchment area was the only independent variable in all 10 linear regression    equations. Predictions of low-flow discharge (80% and 90% exceedance probabilities)    were less accurate than those for the higher flows (Yu and Yang, 2000). A similar    regionalisation scheme to predict FDCs in the United States Mid-Atlantic Region    was developed by Mohamoud (2008). In contrast to the study conducted by Yu and    Yang (2000), a comprehensive set of catchment descriptors (<i>n</i> = 42) were    investigated as potential independent variables in the percentile flow regression    equations. These variables represented catchment land use/land cover, geomorphology,    geology and climate and a step-wise regression approach was used to select the    best predictor variables (Mohamoud, 2008).</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">A major impediment    to the development of FDC region-alisation schemes relates to the number of    gauged rivers in a region. Small sample sizes impact the reliability of equations    to predict FDCs and restrict the number of variables that can be used in the    equations. While this is a problem in many developed countries, the problem    is generally more acute in developing countries where limited resources preclude    installing and maintaining extensive gauging networks. Despite this limitation,    the pressing need for flow information in ungauged catchments requires that    attempts be made to formulate regionalisation schemes using available flow data.</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">Water resource    development and management decisions in South Africa are usually based on monthly    stream flow characteristics (Hughes and Smakhtin, 1996; Smakhtin, 2001). Laws    and policies have been implemented in South Africa that give priority of water    to ecosystems once basic human needs have been met (Acreman and Dunbar, 2004).    The term 'environmental flows' refers to a flow regime which will maintain a    river in some specified condition (Smakhtin, 2007). In many countries, the concept    of minimum flow level was the initial focus for establishing environmental flows,    but now it is increasingly recognised that all elements of a flow regime are    important to manage river ecosystems (Acreman and Dunbar, 2004).</font></p>     ]]></body>
<body><![CDATA[<p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">There is a need    for rapid, low-confidence hydrological predictions in South African catchments    to facilitate initial planning related to ecological instream flow requirements    (Hughes and Hannart, 2003). Synthetic FDCs facilitate planning activities in    ungauged catchments and have become an integral part of environmental flow assessments    (Smakhtin, 2007). This study was initiated to investigate the feasibility of    developing a FDC regionalisation scheme for a critical ecological and economic    region of South Africa, the Cape Floristic Region (CFR). The CFR has a Mediterranean-type    climate (wet winters and dry summers) with catchments that are physi-ographically    and hydromorphometrically distinct from catchments in the summer precipitation    region of South Africa (Seyhan and Hope, 1983). The dominant natural vegetation    is a schlerophyllous shrubland (fynbos) and is home to over 9 000 plant species,    including the highest known concentration of rare species in the world (Cowling    and Hilton-Taylor, 1994; Rouget et al., 2003). The CFR includes the Cape Town    metropolitan area which is dependent on local rivers for most of its water supply.    Catchments in the CFR are under pressure from agricultural development, diminished    stream flow associated with invasion by exotic plant species and rapid urbanisation    (Rouget et al., 2003). These catchments require careful management to ensure    flows can sustain both riparian and estuarine aquatic ecosystems.</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The specific aim    of this study was to develop a monthly FDC regionalisation scheme for small    and intermediate size catchments (area &lt; 300 km<sup>2</sup>) in the CFR.    Larger catchments were excluded since they are more prone to having dams, development    and water abstractions than smaller catchments, and to avoid excessive intra-catchment    heterogeneity. Recognising that the number of gauged catchments for the investigation    was likely to be small, a goal was to design and implement a rigorous regression    approach to predict percentile flows. Since data and vegetation indices (e.g.,    leaf area index, spectral vegetation indices) from the Moderate Resolution Imaging    Spectrometer (MODIS) satellite imagery are readily available, a secondary objective    was to test the utility of MODIS vegetation indices for predicting FDC percentile    flows.</font></p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="3"><b>Methods</b></font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><b>Study region    and catchments</b></font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The CFR covers    87 892 km<sup>2</sup> (<a href="#f1">Fig. 1</a>) and while it is characterised    by a Mediterranean-type climate, a limited amount of precipitation occurs during    the summer months; increasing from west to east across the region (Goldblatt    and Manning, 2002). Mean annual precipitation (MAP) ranges from 200 mm in the    western lowlands to 3 600 mm in the high mountains (Linder, 1991). Free water    evaporation is between 1 250 and 1 600 mm/yr (Seyhan and Hope, 1983). The region    is characterised by diverse physiography which includes sandy coastal plains    underlain by shale; low mountains of limestone, sandstone and conglomerate;    undulating hills underlain by shale located along the inland margins of the    coastal plains; and rugged mountain ranges comprised primarily of sandstones    that rise abruptly to 2 000 m (Goldblatt and Manning, 2002). As mentioned earlier,    the shrubland landscapes of the CFR are characterised by remarkably high species    richness (Linder, 1991; Cowling and Hilton-Taylor, 1994; Rouget et al., 2003).    Forests tend to be located in areas of deeper soils and high precipitation while    most of the hills and valleys are under agriculture (Linder, 1991).</font></p>     <p><a name="f1"></a></p>     <p>&nbsp;</p>     <p align="center"><img src="/img/revistas/wsa/v38n2/04f01.jpg"></p>     <p>&nbsp;</p>     ]]></body>
<body><![CDATA[<p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">River flow data    were obtained from the South African Department Water Affairs (DWA). All gauged    rivers in this region with catchment areas of less than 300 km<sup>2</sup> were    identified as potential candidates for the study. Larger catchments were excluded    in this proof-of-concept study since they are more prone to having dams, development    and water abstractions than smaller catchments. A set of elimination criteria    were applied to identify the final set of study catchments and to minimise uncertainties    in the derivation of percentile flows. Catchments were required to have a minimum    of 10 years of high quality river flow record. In a FDC regionalisation study    in Italy, Castellarin et al. (2007) concluded that 5 years of observed river    flow data was sufficient to obtain consistent estimates of the long-term FDC.    Catchments with impoundments (e.g., dams), water diversions, and significant    urbanisation or agriculture (&gt;5% of catchment area) were excluded and it    was assumed that no major changes in land-cover occurred during the period of    investigation.</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><b>Catchment variables</b></font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">From an initial    pool of 125 catchments, 30 were found to be suitable for the investigation.    Characteristics of these catchments are summarised in <a href="/img/revistas/wsa/v38n2/04t01.jpg">Table    1</a> and their locations in the CFR are indicated in <a href="#f1">Fig. 1</a>.    This small sample size is not unusual for regionalisation studies such as those    conducted by Yu and Yang (2000) in southern Taiwan (n = 10) and Mohamoud (2008)    in the Mid-Atlantic Region, USA (n = 29). Monthly river flow was expressed as    a depth and period-of-record FDCs were constructed for each catchment. These    FDCs were used to determine 11 percentile flows for regionalisation (i.e., Q5,    Q10, Q20, Q30, Q40, Q50, Q60, Q70, Q80, Q90 and Q95).</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The river flow    characteristics and record length for the 30 study catchments are summarised    in <a href="#t2">Table 2</a>. The catchment sample covered a wide range of wetness    conditions, with mean annual river flow (MAQ) ranging from 12 mm for the Sand    River to 1 457 mm for the Wit River (<a href="#t2">Table 2</a>). Percentile    flows for high (Q5), medium (Q50) and low (Q95) flows in <a href="#t2">Table    2</a> also indicate a good distribution of flow regimes in the selected catchments    and two of the rivers (Sand and Smalblaar) are ephemeral.</font></p>     <p><a name="t2"></a></p>     <p>&nbsp;</p>     <p align="center"><img src="/img/revistas/wsa/v38n2/04t02.jpg"></p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">Catchment variables    characterising vegetation, physiography, soils and precipitation were tested    as independent variables in regional regression equations to predict percentile    flows and MAQ. Vegetation descriptors were based on MODIS satellite data from    the Terra satellite which was launched by the US National Aeronautics and Space    Administration (NASA) in 1999. These data are converted on a systematic basis    into derived terrestrial products, including indices that quantify vegetation    cover (Justice et al., 2002). The USGS Land Processes (LP) Distributed Active    Archive Center (DAAC) at the Earth Resources Observation and Science (EROS)    Data Center distributes these MODIS products. Three MODIS vegetation products    were used in the study - 2 spectral vegetation indices and leaf area index (LAI).    The 2 spectral vegetation indices were the Normalised Difference Vegetation    Index (NDVI) and the Enhanced Vegetation Index (EVI). These indices are obtained    from:</font></p>     <p align="center"><img src="/img/revistas/wsa/v38n2/04x01.jpg"></p>     ]]></body>
<body><![CDATA[<p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">where:</font></p>     <blockquote>        <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">p is surface      red or near infrared (nir) reflectance (Tucker, 1979), and</font></p> </blockquote>     <p align="center"><img src="/img/revistas/wsa/v38n2/04x02.jpg"></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"> where:</font></p>     <blockquote>        <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><i>G</i> is a      gain factor</font></p>       <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><i>L</i> is a      canopy background adjustment term (addresses non- linearity of radiation transfer)</font></p>       <p><font  size="2">&#961;</font><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><sub>blue</sub>      is the surface blue reflectance</font></p>       <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">C<sub>1</sub>      and C<sub>2</sub> are weights to correct for different atmospheric aerosol      concentrations (Huete et al., 2002)</font></p> </blockquote>     ]]></body>
<body><![CDATA[<p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The EVI is intended    to be less sensitive to variations in atmospheric conditions than the NDVI and    to have a lesser tendency to saturate at high LAI values (Guo et al., 2007).    LAI is estimated by inversion of a radiative transfer model which uses MODIS    spectral reflectance data (Myneni et al., 2002).</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The NDVI, EVI,    and LAI products for the study area were obtained from the EROS Data Center    DAAC for a regional study funded by the United States National Aeronautics and    Space Administration (Hope et al., 2005) and covered the period April 2000 through    March 2006. The data have a ground resolution of 1 km and values are provided    at 16-day intervals for NDVI and EVI and every 8 days for LAI. Average NDVI,    EVI, and LAI values were calculated for each catchment over the 6-year period.</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">Three physiographic    variables were calculated for each catchment using a 90 m digital elevation    model for the region that was developed using data from the Shuttle Radar Topographic    Mission (SRTM) and provided by the South African Agricultural Research Council,    Institute for Soil Climate and Water (ARC-ISCW). These physiographic variables    were mean elevation, mean slope and drainage density. Data from ARC-ISWC were    used to determine the average fraction of sand, silt and clay for soils in the    catchments. MAP was obtained from 1-km gridded precipitation data also provided    by the ARC-ISCW. Gridded values were based on interpolating data from all available    precipitation gauges in the region and included an adjustment for elevation    (J. Malherbe, 2006). The list of independent variables with their abbreviations    and units of measurement are given in <a href="#t3">Table 3</a>.</font></p>     <p><a name="t3"></a></p>     <p>&nbsp;</p>     <p align="center"><img src="/img/revistas/wsa/v38n2/04t03.jpg"></p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><b>Regression models</b></font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">Multiple regression    models may be developed to produce the single 'best' model for prediction or    to infer causal influences of selected independent variables on the dependent    variable (Mac Nally, 2000). Stepwise regression techniques are often used to    identify predictive models, but there is wide recognition that this is a flawed    approach that is likely to yield spurious results (Mac Nally, 2000). The technique    frequently does not choose the best model predictors and is prone to producing    inflated coefficient of determination values (<i>R</i><sup>2</sup>), leading    to poor model performance in validation (Keith, 2006).</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">Given current computer    power, it is now generally feasible to conduct an exhaustive search of all possible    independent variable combinations ('all-models') to identify the best model    (Mac Nally, 2000). Model selection criteria need to be defined to provide a    compromise between model 'fit' and model 'com-plexity' (Mac Nally, 2000). Model    fit is usually evaluated by an objective function based on the residual sum    of squares while model complexity is indicated by the number of model terms.</font></p>     ]]></body>
<body><![CDATA[<p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The all-models    approach was adopted in this study with a set of catchment descriptors (independent    variables) to identify the best predictive models for the 11 percentile flows    and MAQ. Additive and multiplicative model structures were tested along with    an exhaustive search using untransformed and log-transformed variables to allow    for linear and nonlinear relationships between dependent and independent variables    (Berger, 2004). Given the sample size for model development (<i>n</i> = 30),    model complexity was limited to 4 terms. A 2-step strategy was implemented to    select the best model for each level of complexity. The first step screened    all models for multicolinearity and the remaining models were then ranked according    to their adjusted <i>R</i><sup>2</sup> to identify the best model. Adjusted    <i>R</i><sup>2</sup> is a modified version of <i>R</i><sup>2</sup> which decreases    the <i>R</i><sup>2</sup> value based on the number of explanatory terms in the    model.</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The degree of multi-colinearity    was quantified in Step 1 using the condition index (CI) which is given by:</font></p>     <p align="center"><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><img src="/img/revistas/wsa/v38n2/04x03.jpg"></font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">where:</font></p>     <blockquote>        <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">for a given set      of independent variables, </font><font  size="2">&#955;</font><font face="Verdana, Arial, Helvetica, sans-serif" size="2">      are the eigenvalues of the rescaled crossproduct X'X matrix (Belsley et al.,      1980).</font></p> </blockquote>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The index value    increases with increasing colinearity and, since the index is considered situational,    only rules of thumb exist to reject models. Models with CI greater than 15 were    rejected since Belsley et al. (1980) suggest that weak dependencies are associated    with CI values around 5 or 10 and strong relations are associated with values    above 30.</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The selection of    the best model structure at each percentile flow from the 4 calibrated models    (1 to 4 terms) depends on how well the models predict percentile flows in 'ungauged'    or validation catchments. Unfortunately, a small sample size is often the reality    investigators have to face when they conduct regionalisation studies using gauged    catchment data. The challenge for regression analyses is to have an adequate    sample for model development and validation. The holdout method, which splits    the samples into independent calibration and validation datasets, is hindered    by an inefficient use of sample data in the calibration model which increases    prediction bias (Kohavi, 1995; Blum et al., 1999). A cross-validation approach,    such as the k-fold technique (Kohavi, 1995), can be used to sample all of the    data during calibration when it is not practical to withhold a sub-set for validation.</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The <i>k</i>-fold    validation technique divides the sample data evenly into <i>k</i> groups or    folds, which are then systematically removed from the calibration data as a    validation set. This process is repeated <i>k</i> times, and when <i>k</i> equals    the number of samples in the data the technique is commonly referred to as a    jackknife. Breiman and Spector (1992) and Kohavi (1995) recommend the use of    a 10-fold cross-validation. Although predictions using the 10-fold cross-validation    are generally more biased than in the jackknife approach, prediction variance    is considerably reduced, leading to more accurate estimations and better results    than the jackknife approach (Kohavi, 1995).</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">For each regression    model (1 to 4 terms) developed using all catchments in calibration, we used    a 10-fold cross-validation with the removal of 3 catchments during each resample.    The 3 catchments were selected by stratified random sampling, with 3 strata    defined by magnitude of the percentile flow (3 equal class widths). A random    catchment was selected from the upper-, middle- and lower-flow class for model    validation. Observed and predicted flow values for the 3 validation catchments    from all of the 10-folds were then pooled to evaluate the validation performance    of each model. The Nash and Sutcliffe (1970) coefficient of efficiency (NSE)    was calculated from the observed and estimated flow values to quantify the validation    performance of the 4 models tested for each percentile flow (i.e., a measure    of the overall agreement between observed and predicted flow values for the    validation catchments). The NSE is the ratio of model error variance to the    variance of observed values, subtracted from 1.0. This index was used by Castellarin    et al. (2004) to compare modelled and observed per-centile flows and they considered    values above 0.75 to indicate 'good' agreement.</font></p>     ]]></body>
<body><![CDATA[<p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The single best    model for each percentile flow was selected from the 4 models (1 to 4 terms)    using the largest NSE values determined from the 10-fold validation. We calculated    the relative root-mean-square-error (RRMSE) for each of these regional models    to assess the magnitude of predictive uncertainty. The RRMSE is the root-mean-square-error    divided by the average percentile flow for all catchments.</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><b>Contribution    of MODIS variables</b></font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The regression    approach outlined above was repeated with the 3 MODIS variables excluded from    the pool of potential predictor variables. Validation results (NSE, RRMSE) from    both analyses were compared to assess how these variables affected predictive    uncertainty. We also compared the relative performance of the predictive equations    in each catchment using 'relative error' (RE) as suggested by Castellarin et    al. (2004) and referred to as BIAS by Croker et al (2003). This quantity is    obtained from:</font></p>     <p align="center"><img src="/img/revistas/wsa/v38n2/04x04.jpg"></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">where:</font></p>     <blockquote>        <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><i>Q<sub>E</sub></i>      and <i>Q<sub>0</sub></i> are respectively estimated and observed percen-tile      flows (or MAQ).</font></p> </blockquote>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">For each catchment,    we summed the absolute RE values from each predictive equation used in validation    to quantify the overall uncertainty in the estimated FDC.</font></p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="3"><b>Results and    discussion</b></font></p>     ]]></body>
<body><![CDATA[<p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"> <b>Model selection    and validation</b></font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">Adjusted <i>R</i><sup>2</sup>    values for the 4 best models (1 to 4 variable models) to predict percentile    flows and MAQ are displayed in <a href="#f2">Fig. 2</a>. In most cases, inclusion    of more independent variables increased the adjusted <i>R</i><sup>2</sup> even    though this statistic adjusts for the number of independent variables. Models    for the higher flows (Q5 - Q50) were better than those for the flows below Q60,    with the best adjusted <i>R</i><sup>2</sup> values all greater than 0.8 (<a href="#f2">Fig.    2</a>). The 1-term models had substantially smaller adjusted <i>R</i><sup>2</sup>    values than the other models while there was little difference in the adjusted    <i>R</i><sup>2</sup> of the 3- and 4-term models.</font></p>     <p><a name="f2"></a></p>     <p>&nbsp;</p>     <p align="center"><img src="/img/revistas/wsa/v38n2/04f02.jpg"></p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The NSE values    obtained using 10-fold validation of the best models are given in <a href="#f3">Fig.    3</a>. As expected from the model development results, prediction of percentile    flows above Q60 were notably better than predictions for the low flows (<a href="#f3">Fig.    3</a>). However, the model performance did not improve consistently with the    number of variables included in the equations as was the case in the model development    phase. Models with 4 variables were not the best models in validation for any    of the percentile flows or MAQ (<a href="#f3">Fig. 3</a>). Instead, models with    3 variables were the best models in validation except for Q50 (2 variables)    and Q70 (1 variable). Only equations for percen-tile flows above Q50 and for    MAQ had NSE values greater than 0.75, the threshold suggested by Castellarin    et al. (2004) to indicate 'good' FDC models.</font></p>     <p><a name="f3"></a></p>     <p>&nbsp;</p>     <p align="center"><img src="/img/revistas/wsa/v38n2/04f03.jpg"></p>     ]]></body>
<body><![CDATA[<p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The best regression    models for each percentile flow and MAQ based on validation performance are    given in <a href="/img/revistas/wsa/v38n2/04t04.jpg">Table 4</a> along with their adjusted <i>R</i><sup>2</sup>    and CI values from model development and the validation statistics (NSE, RRMSE).    In all cases, additive models were selected over the multiplicative models.    Models for percentile flows from Q5 to Q50 and for Q70 and MAQ were linear while    the remaining models for low flows included logarithmic terms (<a href="/img/revistas/wsa/v38n2/04t04.jpg">Table    4</a>). All models included MAP except the Q70 and Q80 models, while soil clay    fraction (CLAY) appeared in the high-flow models and in the MAQ and Q95 models.    Although the adjusted <i>R</i><sup>2</sup> and NSE values from model development    and validation were greater than 0.7 for percentile flows Q5 to Q50, validation    RRMSE values were all greater than 37% (<a href="/img/revistas/wsa/v38n2/04t04.jpg">Table 4</a>),    indicating potentially large uncertainty in predicting these quantities in ungauged    catchments.</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">Since the prediction    of low flows was found to be uncer-tain, it seemed likely that the overall regionalisation    approach may be better suited to wet, rather than dry catchments. For each catchment,    we summed the absolute RE values from each predictive equation used in validation    and then plotted this total RE against MAP (<a href="#f4">Fig. 4</a>). The upper    limit of total RE in <a href="#f4">Fig. 4</a> (broken line) increased as MAP    decreased, indicating the potential for larger uncertainties in the drier catchments.    The total absolute RE for the driest catchment (Sand River) was considerably    larger than values in the other catchments, indicating a possible threshold    MAP (less than 300 mm) for appropriate use of this approach. These findings    are similar to those reported by Yu and Yang (2000) and Hope and Bart (2012),    who also found weaker models for predicting the low percentile flows for FDCs    in Taiwan and southern California USA, respectively. In each of these studies,    processes controlling low flows may not have been adequately represented by    the variables used in the models or uncertainties in the measurement of low    flows may have contributed to predictive errors.</font></p>     <p><a name="f4"></a></p>     <p>&nbsp;</p>     <p align="center"><img src="/img/revistas/wsa/v38n2/04f04.jpg"></p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><b>Effect of MODIS    variables</b></font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">When MODIS variables    were included in the development phase of the flow prediction models, 7 of the    12 equations included these variables (<a href="/img/revistas/wsa/v38n2/04t04.jpg">Table 4</a>).    The EVI was included in equations for Q5, Q20, Q30, Q40 and MAQ while the NDVI    was in the equation for Q10 and LAI was selected for the Q80 model. The 10-fold    validation results (NSE, RRMSE) given in <a href="#f5">Fig. 5</a> are for equations    developed with MODIS variables excluded from the model development phase. All    NSE values in <a href="#f5">Fig. 5</a> were smaller than values for corresponding    models that did include MODIS variables (<a href="#f3">Fig. 3</a>).</font></p>     <p><a name="f5"></a></p>     ]]></body>
<body><![CDATA[<p>&nbsp;</p>     <p align="center"><img src="/img/revistas/wsa/v38n2/04f05.jpg"></p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">The best regression    models without MODIS variables that were identified using the 10-fold validation    are given in <a href="/img/revistas/wsa/v38n2/04t05.jpg">Table 5</a>. While models to predict    percentile flows that included MODIS variables (<a href="/img/revistas/wsa/v38n2/04t04.jpg">Table    4</a>) had better calibration results than models without MODIS variables (<a href="/img/revistas/wsa/v38n2/04t05.jpg">Table    5</a>), the differences in NSE and RRMSE were not substantial. At most, the    RRMSE was 5% smaller with MODIS variables in the equations. Except for Q70 and    Q80, catchment MAP was the first variable to enter the regression equations,    reflecting the dominant effect of this variable for the estimation of percentile    flows and MAQ. While these results appear to indicate that vegetation has a    small effect on river flows, the results also may be a consequence of the research    methodology. The vegetation indices were represented in the regression equations    as area- and time-averaged values for each catchment. Vegetation effects on    hydrological fluxes in different parts of the catchments and at different times    of the year could not be represented. For example, transpiration associated    with phreatophytic vegetation located within the 0.9 0.8 0.7 0.6 0.5 0.4 0.3    0.2 0.10 riparian zone could be expected to impact low flows more than transpiration    from vegetation on the hill slopes. This was demonstrated by Hope et al. (2009),    who found a significant relationship between a spectral vegetation index (NDVI)    in the lowland area of a CFR catchment (Molenaars) and annual flow volume, but    no relationship when the index was calculated for the upland areas.</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">To assess the effect    of MODIS variables on the prediction of percentile flows in individual catchments,    the average RE values from validation of models with and without MODIS variables    were plotted against each other (<a href="#f6ad">Figs. 6a-6g</a>). The exclusion    of MODIS variables from the predictive equations had little effect on predictive    accuracy in most catchments, with most of the points plotting around the 1:1    lines (<a href="#f6ad">Fig. 6</a>). However, in 3 catchments (Sand River, Brandwag    River and Wilge River) the exclusion of MODIS variables caused large predictive    errors for percentile flows Q5-Q40 and MAQ (<a href="#f6ad">Fig. 6</a>). Prediction    errors in the Smalblaar River catchment were also notably larger for models    without MODIS variables to predict Q40 (<a href="#f6be">Fig. 6e</a>) and MAQ    (<a href="#f6g">Fig. 6g</a>), but smaller to predict Q80 (<a href="#f6cf">Fig.    6f</a>). These 4 catchments (Sand, Brandwag, Wilge and Smalblaar Rivers) are    also the catchments with the 4 lowest mean annual flows (MAQ) (<a href="#t2">Table    2</a>). Since soil moisture rather than the amount of vegetation tends to be    the controlling variable affecting evaporative losses and river flow in water-limited    catchments, it is not apparent why vegetation indices were more important variables    in these drier catchments than in wetter catchments. However, given the small    sample of catchments used in this study, direct conclusions regarding the effect    of vegetation on river flows may not be appropriate.</font></p>     <p><a name="f6ad"></a></p>     <p>&nbsp;</p>     <p align="center"><img src="/img/revistas/wsa/v38n2/04f06ad.jpg">    <br>   <a name="f6be"></a> <img src="/img/revistas/wsa/v38n2/04f06be.jpg">    <br>   <a name="f6cf"></a> <img src="/img/revistas/wsa/v38n2/04f06cf.jpg">    ]]></body>
<body><![CDATA[<br>   <a name="f6g"></a> <img src="/img/revistas/wsa/v38n2/04f06g.jpg"></p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="3"><b>Conclusion</b></font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">A goal of this    study was to develop regional regression equations that could be used to predict    monthly FDC percentile flows and MAQ from readily measureable catchment variables,    including satellite-derived vegetation indices. The all-models approach with    10-fold validation was found to be suitable for use with the restricted catchment    sample size available for this study. The challenge of predicting low flows    in semi-arid catchments is well documented (e.g., Pilgrim et al., 1988; Croke    and Jakeman, 2008) and, as expected, the prediction of the larger percentile    flows (Q5 - Q50) and MAQ were notably better than prediction of low flows. Based    on the prediction equations for FDC percentile flows in the CFR, it may be concluded    that catchment mean annual precipitation, physiography and soils were more important    predictive variables than MODIS vegetation indices. Use of MODIS vegetation    variables in the regression equations did not result in substantially better    calibration results in most catchments, but did reduce predictive uncertainty    substantially in 3 or 4 catchments depending on the flow calculation.</font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">While the validation    results of this study pointed to large uncertainties in the prediction of percentile    flows, the set of equations provide a means for rapid, low-confidence estimates    for initial catchment planning, as suggested by Hughes and Hannart (2003). Uncertainties    in the measurement of river flows, due to flows exceeding the available rating    curves and the possibility that some low flows were impacted by undocumented    abstractions, may have contributed to prediction uncertainties. More reliable    synthetic FDCs may be attainable if a larger sample of study catchments could    be identified. Alternative approaches should also be investigated where regression    equations are used to derive parameters of mathematical or probabilistic FDC    models (e.g., Niadas, 2005; Castellarin et al., 2007).</font></p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="3"><b>Acknowledgements</b></font></p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">Assistance with    data processing and the construction of the tables and figures was provided    by Noah Albers and Keith James (San Diego State University). We gratefully acknowledge    the assistance with data sets provided by the Agricultural Research Council,    Institute for Soil Climate and Water (Terrence Newby, Talita Germishuyse, Johan    Malherbe and Ian Kotze). This study was funded by the U.S. National Aeronautics    and Space Administration, Land Cover Land Use Change Program, Grant No. NNG05GR14G.</font></p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="3"><b>References</b></font></p>     ]]></body>
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Sci. 15</i> 323-331.</font>&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;[&#160;<a href="javascript:void(0);" onclick="javascript: window.open('/scielo.php?script=sci_nlinks&ref=941820&pid=S1816-7950201200020000400036&lng=','','width=640,height=500,resizable=yes,scrollbars=1,menubar=yes,');">Links</a>&#160;]<!-- end-ref --><!-- ref --><p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">YU PS and YANG    TC (2000) Using synthetic flow duration curves for rainfall-runoff model calibration    at ungauged sites. <i>Hydrol. Process. 14</i> 117-133.</font>&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;&nbsp;[&#160;<a href="javascript:void(0);" onclick="javascript: window.open('/scielo.php?script=sci_nlinks&ref=941821&pid=S1816-7950201200020000400037&lng=','','width=640,height=500,resizable=yes,scrollbars=1,menubar=yes,');">Links</a>&#160;]<!-- end-ref --><p>&nbsp;</p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2">Received 11 July    2011;    ]]></body>
<body><![CDATA[<br>   accepted in revised form 2 April 2012.</font></p>     <p>&nbsp;</p>     <p>&nbsp;</p>     <p><font face="Verdana, Arial, Helvetica, sans-serif" size="2"><a name="back"></a><a href="#top">*</a>    To whom all correspondence should be addressed. 619-594-2777; fax: 619-594-4938;    e-mail: <a href="mailto:hope1@mail.sdsu.edu">hope1@mail.sdsu.edu</a></font></p>      ]]></body>
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