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African Natural History

On-line version ISSN 2305-7963
Print version ISSN 1816-8396

Afr. nat. history (Online) vol.6  Cape Town Jan. 2010


Cretaceous faunas from Zululand and Natal, South Africa. The ammonite subfamily Acanthoceratinae de Grossouvre, 1894



William James KennedyI; Herbert Christian KlingerII

IOxford University Museum of Natural History. Parks Road, Oxford. 0X1 3PW, U.K. E-mail:
IINatural History Collections Department, Iziko South African Museum. P.O. Box 61, Cape Town, 8000 South Africa and Department of Geological Sciences, University of Cape Town, Private Bag, Rondebosch, 7701 South Africa. E-mail:




Ammonites of the subfamily Acanthoceratinae occur in abundance at a single locality in KwaZulu, the eastern part of the Skoenberg, on the lower reaches of the Mzinene River, north-northeast of Hluhluwe. This is the classic locality from which William Anderson collected the material described by G.C. Crick in 1907 as being from the 'deposit at the north end of False Bay.' The fossils occur weathered out on the surface; there is no section. The ammonites described below are predominantly Middle Cenomanian, with indications of the presence of lower Upper Cenomanian on the basis of records elsewhere. The following species are described: Acanthoceras flexuosum Crick, 1907 (of which Acanthoceras crassiornatum Crick, 1907, Acanthoceras munitum Crick, 1907, Acanthoceras expansum Crick, 1907, Acanthoceras robustum Crick, 1907, Acanthoceras quadratum Crick, 1907, Acanthoceras hippocastanum Crick, non J. de C. Sowerby, and Acanthoceras latum Crick, 1907, are synonyms), Acanthoceras cornigerum Crick, 1907, Protacanthoceras subwaterloti (Venzo, 1936), Cunningtonicerasl sp. a and sp. b, Calycoceras (Gentoniceras ) gentoni (Brongniart, 1822) (of which Acanthoceras paucinodatum Crick, 1907 is a synonym), Calycoceras (Newbokiiceras) asiaticum asiaticum (Jimbo, 1894), Calycoceras (Newboldiceras) asiaticum spinosum (Kossmat, 1897), Calycoceras (Newboldiceras) cf. asiaticum hunteri (Kossmat, 1897), Calycoceras [Newboldiceras) planecostatum (Kossmat, 1897), Calycoceras {Newboldiceras) laticostatum (Crick, 1907), Calycoceras (Newboldiceras) breistrofteri (Collignon, 1937) and Pseudocalycoceras harpax (Stoliczka, 1864).

Key words: ammonites, Acanthoceratinae, Cenomanian, Mzinene Formation, KwaZulu- Natal, South Africa.



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Kennedy thanks the staff of the Department of Earth Sciences, Oxford, and the University Museum of Natural History, Oxford, for technical support. The financial assistance of the Strakosh Bequest and the Oppenheimer Fund (Oxford) is acknowledged. Lorna Steel (The Natural History Museum, London) provided access to specimens in her care. Klinger acknowledges the financial aid of the NRF (South Africa) and technical staff of the Natural History Collections Department, Iziko South African Museum. The originals of Figs 1-6, 30, 31 were supplied by the photographic unit of the Natural History Museum, London.



AMÈDRO, F., 1994. In: ROBASZYNSKI, F., CARON, M., AMÈDRO, F., DUPUIS, C., HARDENBOL, J., GONZALEZ DONOSO, J.M., LINARES, D. & GARTNER, S. Le Cénomanien de la région de KalatSenan (Tunisie centrale): litho-biostratigraphie et interprétation séquentielle. Revue de Paléobiologie 12 : 351-505.         [ Links ]

ANDERSON, W. 1907. Third and Final Report of the Geological Survey of Natal and Zululand. 299 pp. London: West, Newman & Co.         [ Links ]

AVNIMELECH, M.A. & SHORESH, R. 1962. Les céphalopodes cénomaniens des environs de Jerusalem. Bulletin de la Société Géologique de la France (7)4: 528-535.         [ Links ]

BASSE, E. 1931. Monographie paléontologique du Crétacé de la Province de Maintirano. Mémoires Géologiques du Service des Mines de Madagascar (1931): 1-86 .         [ Links ]

BASSE, E. 1940. Les céphalopodes crétacés des massifs côtiers syriens, pt. 2. Notes et Mémoires du Haute Commissariat de la République Française en Syrie et au Liban, 2, études Paléontologiques: 411-472.         [ Links ]

BESAIRIE, H. 1930. Les rapports du Crétacé malgache avec le Crétacé de l'Afrique australe. Bulletin de la Société Géologique de France (4)30: 613-643.         [ Links ]

BOSC, L.A.G. 1801. (An. 13). In: ROISSY, F. Histoire Naturelle générale et particulière, des Mollusques, Animaux sans vertèbres et à sang blanc. Ouevrage faisant suite à aux oeuvres de Leclerc de Buffon, et partie du Cours complet d'Histoire naturelle, rédigé par C.S. Sonnini, membre de plusieurs Sociétés savantes. Continué par. F. de Roissy. viii + 448 pp. Paris.         [ Links ]

BRONGNIART, A. 1822. Sur quelques terrains de Craie hors du Bassin de Paris. In: G. CUVIER & A. BRONGNIART, Description géologique des environs de Paris, 3rd edn. 1-428. Paris: Dufour et D'Ocagne,         [ Links ].

COBBAN, W.A., HOOK, S.C. & KENNEDY, WJ. 1989. Upper Cretaceous rocks and ammonite faunas of south-western New Mexico. Memoir of the New Mexico Bureau of Mines and Mineral Resources 45: 1-137.         [ Links ]

COLLIGNON, M. 1937. Ammonites Cénomaniennes du sud-ouest de Madagascar. Annales géologiques du Service des Mines de Madagascar 8: 31-69.         [ Links ]

COLLIGNON, M. 1939. Fossiles cénomaniens et turoniens du Menabe (Madagascar). Annales géologiques du Service des Mines de Madagascar 10: 59-105.         [ Links ]

COLLIGNON, M. 1964. Atlas des fossiles caractéristiques de Madagascar (Ammonites), XI. Cénomanien. xi + 1-152. Tananarive: Service Géologique.         [ Links ]

COLLIGNON, M., 1967. Les céphalopodes crétacés du bassin côtier de Tarfaya. Notes et Mémoires du Service géologique du Maroc 175: 1-149 (1966 imprint).         [ Links ]

CRICK, G.C. 1907. Cretaceous fossils of Natal. In: ANDERSON, W. Third and Final Report of the Geological SurveyofNatal and Zululand, pp. 161-250. London: West, Newman & Co.         [ Links ]

DIETRICH, W.O. 1938. Zur Stratigraphie der Kreide im nördlichen Zululand. Zentralblatt für Mineralogie, Geologie und Paläontologie (B) 1938: 228-240.         [ Links ]

DU TOIT, A.L. & HOEPEN, E.C.N. VAN 1929. Guide Book. Excursion C18, Durban-Zululand. 1-20. XV International Geological Congress: Pretoria, South Africa.         [ Links ] DOUVILLÉ, R. 1911. Ammonites gentoni Defrance, in Brongniart, 1822. Palaeontologia Universalis n.s.: Fiche 223.         [ Links ]

FÖRSTER, R. 1975. Die geologische Entwicklung von süd-Mozambique seit der Unterkreide und die Ammoniten-Fauna von Unterkreide und Cenoman. Geologisches Jahrbuch B12: 3-324.         [ Links ]

GALE, A.S. 1995. Cyclostratigraphy and correlation of the Cenomanian Stage in Western Europe. Geological Society Special Publication 85: 177-197.         [ Links ]

GROSSOUVRE, A. de 1894. Recherches sur la Craie supérieure, 2, Paléontologie. Les ammonites de la Craie supérieure. Mémoires du Service de la Carte Géologique détaillée de la France. 1-264. (misdated 1893).         [ Links ]

HEINZ, R. 1930. Ueber Kreide-Inoceramen der südafrikanischen Union. Comptes Rendus, XV International Geological Congress, Cape Town 2: 681-687.         [ Links ]

HOEPEN, E.C.N. VAN 1926. Oor die Krytafsettinge van Soeloeland. South African Journal of Science 23: 216-222.         [ Links ]

HOEPEN, E.C.N. VAN 1929. Die Krytfauna van Soeloeland. Paleontologiese Navorsing van die Nasionale Museum, Bloemfontein 1: 1-38.         [ Links ]

HYATT, A. 1900. Cephalopoda. In: ZITTEL, K.A. VON 1896-1900, Textbook of Palaeontology, pp. 502-604, transl. EASTMAN, C.R. London and New York: Macmillan.         [ Links ]

JIMBO, K. 1894. Beiträge zur Kenntniss der Fauna der Kreideformation von Hokkaido. Paläontologische Abhandlungen. (n.s.) 2: 147-194.         [ Links ]

KAPLAN, U., KENNEDY, W.J., LEHMANN, J. & MARCINOWSKI, R. 1998. Stratigraphie und Ammonitenfaunen des westfälischen Cenoman. Geologie und Paläontologie in Westfalen 51: 1-236.         [ Links ]

KENNEDY, WJ. 1971. Cenomanian ammonites from southern England. Special Papers in Palaeontology 8: v + 1-133.         [ Links ]

KENNEDY, W.J. 1978 The middle Cretaceous of Zululand and Natal, South Africa. Annales du Muséum d'Histoire Naturelle de Nice 4: XVIII.1-XVIII.29. (misdated 1976).         [ Links ]

KENNEDY, W J. 1988. Late Cenomanian and Turonian ammonite faunas from north-east and central Texas. Special Papers in Palaeontology 8: 1-133.         [ Links ]

KENNEDY, W J. 1994. Cenomanian ammonites from Cassis, Bouches-du-Rhône, France. Palaeopelagos, Special Volume 1: 209-254.         [ Links ]

KENNEDY, W.J., BILOTTE, M. & HANSOTTE, M. 1966. Cenomanian ammonites from Pech de Foix (Ariège,France). Géobios 29: 307-318.         [ Links ]

KENNEDY, W.J. & JOLKICEV, N. 2004. Middle Cenomanian ammonites from the type section of the Sanandinovo Formation of northern Bulgaria. Acta Geologica Polonica 54: 369-380.         [ Links ]

KENNEDY, W J. & JUIGNET, P. 1994. A revision of the ammonite faunas of the type Cenomanian, 5. Acanthoceratinae (Calycoceras (Calycoceras), C. (Gentoniceras) and C. (Newboldiceras)). Cretaceous Research 15: 17-57.         [ Links ]

KENNEDY, W J. & KLINGER, H.C. 1975. Cretaceous faunas from Zululand and Natal, South Africa. Introduction, Stratigraphy. Bulletin of the British Museum (Natural History) Geology 25: 263-315.         [ Links ]

KORN, D., EBBIGHAUSEN, V., BOCKWINKEL, J. & KLUG, C. 2003. The A-mode ontogeny in prolecanitid ammonites. Palaeontology 46: 1123-1132.         [ Links ]

KOSSMAT, F. 1895-1898. Untersuchungen über die Südindische Kreideformation. Beiträge zur Paläontologie Österreich-Ungarns und des Orients. 9 (1895): 97-203(1-107); 11(1897): 1-46 (108-153); 11 (1898): 89-152 (154-217).         [ Links ]

KULLMANN, J. & WIEDMANN, J. 1970. Significance of sutures in phylogeny of Ammonoidea. University of Kansas, Paleontological Contributions 42: 1-32.         [ Links ]

LAMARCK, J.P.B.A. de M. de 1801. Système des Animaux sans vertèbrès. vii + 1-432. Paris: The author; Deterville.         [ Links ]

LEHMANN, J. & HERBIG, H-G. 2009. Late Cretaceous ammonites the from Bou Angueur syncline (Middle Atlas, Morocco)-stratigraphic and paleobiogeographic implications. Palaeontographica A 289:45-87.         [ Links ]

MANTELL, G.A. 1822. The Fossils of the South Downs; or Illustrations of the Geology of Sussex. xvi + 1-327. London: Lupton Relfe.         [ Links ]

MATSUMOTO, T. & KAWANO, T. 1975. A find of Pseudocalycoceras from Hokkaido. Transactions and Proceedings of the Palaeontological Society of Japan 97: 7-21.         [ Links ]

MATSUMOTO, T., MURAMOTO, T. & TAKAHASHI, T. 1969. Selected acanthoceratids from Hokkaido. Memoirs of the Faculty of Science Kyushu University (D) Geology 19: 251-296.         [ Links ]

MATSUMOTO, T., SAITO, R. & FUKADA, A. 1957. Some acanthoceratids from Hokkaido. Memoirs of the Faculty of Science Kyushu University (D) Geology 6: 1-45.         [ Links ]

MONTANARO, E. & LANG, Z. 1937. Coelenterati, echinodermi, e brachiopodi del cretaceo medio-superiore dello Zululand. Palaeontographia Italica 37:193-201.         [ Links ]

MOREMAN, W.L. 1942. Paleontology of the Eagle Ford group of north and central Texas. Journal of Paleontology 16: 192-220.         [ Links ]

NEUMAYR, M. 1875. Die Ammoniten der Kreide und die Systematik der Ammonitiden. Zeitschrift der Deutschen Geologischen Gesellschaft 27: 854-942.         [ Links ]

PASSY, A. 1832. Description géologique du département de la Seine-inférieure. xvi + 1-371. Rouen: Imprimerie Nicétas Periaux.         [ Links ]

PERVINQUIÈRE, L. 1907. Etudes de paléontologie tunisienne. 1. Céphalopodes des terrains secondaires. Carte Géologique de la tunisie. v + 1-438 . Paris: de Rudeval.         [ Links ]

SHARPE, D. 1853-57. Description of the fossil remains of Mollusca found in the Chalk of England. I, Cephalopoda. Palaeontographical Society Monographs. 1-68. 1-26 (1853); 27-36 (1855); 37-68 (1857).         [ Links ]

SOCIN, C. 1939. Gasteropodi e Lamellibranchi del cretaceo medio-superiore dello Zululand. Palaeontographia Italica 40: 21-38.         [ Links ]

SPATH, L.F. 1923. On the ammonite horizons of the Gault and contiguous deposits. Summary of Progress of the Geological Survey of Great Britain for 1922: 139-149.         [ Links ]

SPATH, L.F. 1926. On new ammonites from the English Chalk. Geological Magazine 63: 77-83.         [ Links ]

STOLICZKA, F. 1863-1866. The fossil cephalopoda of the Cretaceous rocks of southern India. Ammonitidae with revision of the Nautilidae etc. Memoirs of the Geological Survey of India. (1), Palaeontologica Indica, 3: (1), 41-56 (1863); (2-5), 57-106 (1864); (6-9), 107-154 (1865); (10-13), 155-216 (1866).         [ Links ]

TAUBENHAUS, H. 1920. Die Ammoneen der Kreideformation Palästinas und Syriens. Zeitschrift des Deutschen PalästinaVereins 43: 1-58.         [ Links ]

THOMEL, G. 1966. Ammonites. In: PORTHAULT, B., THOMEL, G. & VILLOUTREYS, O. de. Étude biostratigraphique du Cénomanien du bassin supérieur de l'Esteron (Alpes Maritimes). LeproblèmedelalimiteCénomanien-Turoniandanslesud-estde la France. Bulletin de la Société Géologique de France (7)8: 423-439.         [ Links ]

THOMEL, G. 1972. Les Acanthoceratidae cénomaniens des chaines subalpines méridionales. Mémoires de la Société Géologique de France (N.S.) 116: 1-204.         [ Links ]

VENZO, S. 1936. Cefalopodi del Cretaceo medio-superiore dello Zululand. Palaeontographia Italica 36: 59-133 (1-75).         [ Links ]

WILMSEN, M. 1977. Das Oberalb und Cenoman im Nord-kantabrischen Becken (Provinz Kantabrian, Nordspanien): Faziesentwicklung, bio-und Sequenzstratigraphie. Berliner geowissenschaftliche Abhandlungen E23: 1-167.         [ Links ]

WRIGHT, C.W. 1996. In: Treatise on Invertebrate Paleontology Part L, Mollusca 4: Cretaceous Ammonoidea. xx + 1-362 (with Contributions by J.H. Calloman [sic] and M.K. Howarth). Boulder, Colorado and Lawrence, Kansas: Geological Society of America and University of Kansas Press.         [ Links ]

WRIGHT, C.W. & KENNEDY, W J. 1981. The Ammonoidea of the Plenus Marls and the Middle Chalk. Palaeontographical Society Monographs: 1-148.         [ Links ]

WRIGHT, C.W. & KENNEDY, W J. 1987. The Ammonoidea of the Lower Chalk. Part 2. Palaeontographical SocietyMonographs: 127-215.         [ Links ]

WRIGHT, C.W. & KENNEDY, W J. 1990. The Ammonoidea of the Lower Chalk. Part 3. Palaeontographical SocietyMonographs: 219-294.         [ Links ]

ZABORSKI, P.M.P. 1985. Upper Cretaceous ammonites from the Calabar region, southeast Nigeria. Bulletin of the British Museum (Natural History) Geology 39: 1-72.         [ Links ]

ZITTEL, K.A. VON 1884. Handbuch der Palaeontologie 1, Abt. 2; Lief 3, Cephalopoda. pp. 329-522. Munich & Leipzig: R. Oldenbourg.         [ Links ]



Submitted 6 January 2010
Accepted 23 April 2010

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